DGRPool

GWAS results were computed on dm3 genome.

If you want to make an independent analysis with DGRP2 or PLINK2, you can find here prepared input files for this phenotype: DGRP2 results PLINK2 results

Full results

Phenotype distribution and association to known covariates

Shapiro-Wilk test of normality (p-value=5.648e^-23)
Null Hypothesis is rejected (p<=0.05).
Interpretation: NOT NORMAL distribution of this phenotype

We tested for association of this phenotype with 6 known covariates Huang et al., 2014 using both ANOVA and Kruskal-Wallis association tests.

ANOVA test

Covariate	p-value	Significance
In_2R_NS	0.06011

Kruskal-Wallis test
No significant result.

PNG PDF

QQplot

PNG PDF

Manhattan plot

GWAS results

1622 significant hits at p-value < 0.001. The 1000 top results are displayed in the table below.

We performed a gene enrichment calculation (Fisher's exact test) of all GWAS-associated genes at p < 0.001. Genesets were taken from two database: the Gene Ontology (GO) and the FlyBase phenotypes. GO enrichment FlyBase phenotype enrichment

Variant impact HIGH MODERATE LOW MODIFIER

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Chromosome	Position	ID	Reference	Variant	p-value	FDR_BH	Transcript annotation	Binding site annotation
2R	15792957	2R_15792957_SNP [dm3] 2R_19905452_SNP [dm6]	T	C	5.315e^-8	0.1	INTERGENIC
3L	3094996	3L_3094996_SNP [dm3] 3L_3094996_SNP [dm6]	A	C	4.394e^-7	0.412	UPSTREAM tRNA:CR32288 & 1 other INTRON Cht7	TF binding site mE1_TFBS_cad
2R	15329231	2R_15329231_SNP [dm3] 2R_19441726_SNP [dm6]	A	C	1.361e^-6	0.706	SYNONYMOUS_CODING Spt5 UPSTREAM RpL11	TF binding site BDTNP1_TFBS_dl mE1_TFBS_HSA
3L	4176724	3L_4176724_SNP [dm3] 3L_4176724_SNP [dm6]	G	T	1.506e^-6	0.706	SYNONYMOUS_CODING Teh4 INTRON CG18675
3L	3095378	3L_3095378_SNP [dm3] 3L_3095378_SNP [dm6]	T	A	1.978e^-6	0.735	UPSTREAM tRNA:CR32288 & 2 others INTRON Cht7
3L	3096577	3L_3096577_SNP [dm3] 3L_3096577_SNP [dm6]	G	T	2.484e^-6	0.735	UPSTREAM tRNA:CR32285 & 1 other DOWNSTREAM tRNA:CR32288 & 3 others INTRON Cht7	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_Med mE1_TFBS_HSA
3L	3096817	3L_3096817_SNP [dm3] 3L_3096817_SNP [dm6]	T	C	3.452e^-6	0.735	UPSTREAM tRNA:CR32286 DOWNSTREAM tRNA:CR32272 & 3 others INTRON Cht7	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_Med mE1_TFBS_HSA
3L	3096219	3L_3096219_SNP [dm3] 3L_3096219_SNP [dm6]	T	A	3.474e^-6	0.735	UPSTREAM tRNA:CR32287 & 1 other DOWNSTREAM tRNA:CR32288 & 2 others INTRON Cht7	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_Med mE1_TFBS_HSA
3R	15358868	3R_15358868_SNP [dm3] 3R_19533146_SNP [dm6]	A	C	3.981e^-6	0.735	INTRON Dys	TF binding site mE1_TFBS_inv
2L	13123705	2L_13123705_SNP [dm3] 2L_13123705_SNP [dm6]	G	T	4.656e^-6	0.735	INTERGENIC
3L	3096214	3L_3096214_SNP [dm3] 3L_3096214_SNP [dm6]	A	G	5.216e^-6	0.735	UPSTREAM tRNA:CR32287 & 1 other DOWNSTREAM tRNA:CR32288 & 2 others INTRON Cht7	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_Med mE1_TFBS_HSA
2L	10557769	2L_10557769_SNP [dm3] 2L_10557769_SNP [dm6]	C	A	5.357e^-6	0.735	INTRON Trim9	TF binding site mE1_TFBS_HSA mE1_TFBS_inv
3R	15358879	3R_15358879_SNP [dm3] 3R_19533157_SNP [dm6]	T	A	5.364e^-6	0.735	INTRON Dys	TF binding site mE1_TFBS_inv
3R	15358867	3R_15358867_SNP [dm3] 3R_19533145_SNP [dm6]	G	T	5.489e^-6	0.735	INTRON Dys	TF binding site mE1_TFBS_inv
3L	3096971	3L_3096971_SNP [dm3] 3L_3096971_SNP [dm6]	T	A	6.282e^-6	0.786	UPSTREAM tRNA:CR32286 & 1 other DOWNSTREAM tRNA:CR32289 & 2 others INTRON Cht7	TF binding site BDTNP1_TFBS_Med mE1_TFBS_HSA
2L	1102011	2L_1102011_SNP [dm3] 2L_1102011_SNP [dm6]	G	A	7.050e^-6	0.811	INTRON CG4629
3L	4176703	3L_4176703_SNP [dm3] 3L_4176703_SNP [dm6]	G	T	7.518e^-6	0.811	SYNONYMOUS_CODING Teh4 INTRON CG18675
3L	11661243	3L_11661243_SNP [dm3] 3L_11668143_SNP [dm6]	A	G	7.780e^-6	0.811	INTRON CG32085
3R	4602693	3R_4602693_SNP [dm3] 3R_8776971_SNP [dm6]	C	A	8.681e^-6	0.816	SYNONYMOUS_CODING Pif1B UTR_3_PRIME Pif1A	TF binding site mE1_TFBS_HSA
2R	15828470	2R_15828470_SNP [dm3] 2R_19940965_SNP [dm6]	G	A	9.052e^-6	0.816	UPSTREAM CR43421 INTERGENIC	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_prd BDTNP1_TFBS_twi mE1_TFBS_HSA
2R	20110237	2R_20110237_SNP [dm3] 2R_24222714_SNP [dm6]	G	C	9.860e^-6	0.816	INTRON CG30419
2R	20115290	2R_20115290_SNP [dm3] 2R_24227767_SNP [dm6]	C	A	1.030e^-5	0.816	INTRON CG30419
3L	3096973	3L_3096973_SNP [dm3] 3L_3096973_SNP [dm6]	G	A	1.032e^-5	0.816	UPSTREAM tRNA:CR32286 & 1 other DOWNSTREAM tRNA:CR32289 & 2 others INTRON Cht7	TF binding site BDTNP1_TFBS_Med mE1_TFBS_HSA
X	2703387	X_2703387_SNP [dm3] X_2809354_SNP [dm6]	C	A	1.050e^-5	0.816	UPSTREAM CG12498 INTRON kirre
2R	1342713	2R_1342713_INS [dm3] 2R_5455208_INS [dm6]	T	TAAGATTAAATTGTAGTTTG...	1.087e^-5	0.816	UTR_3_PRIME laccase2 DOWNSTREAM laccase2
3L	3097469	3L_3097469_SNP [dm3] 3L_3097469_SNP [dm6]	T	C	1.209e^-5	0.858	UPSTREAM tRNA:CR32286 & 1 other DOWNSTREAM tRNA:CR32285 INTRON Cht7
3R	15358853	3R_15358853_SNP [dm3] 3R_19533131_SNP [dm6]	A	C	1.328e^-5	0.858	INTRON Dys
2L	16078700	2L_16078700_SNP [dm3] 2L_16078700_SNP [dm6]	T	A	1.403e^-5	0.858	INTRON beat-Ia
2R	9924405	2R_9924405_SNP [dm3] 2R_14036900_SNP [dm6]	C	A	1.421e^-5	0.858	INTRON mam	TF binding site BDTNP1_TFBS_dl mE1_TFBS_HSA
3R	15358855	3R_15358855_SNP [dm3] 3R_19533133_SNP [dm6]	A	C	1.455e^-5	0.858	INTRON Dys
3R	15358852	3R_15358852_SNP [dm3] 3R_19533130_SNP [dm6]	G	C	1.468e^-5	0.858	INTRON Dys
3R	10680426	3R_10680426_SNP [dm3] 3R_14854704_SNP [dm6]	T	A	1.574e^-5	0.858	UPSTREAM btsz INTRON btsz	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_Med BDTNP1_TFBS_sna BDTNP1_TFBS_twi mE1_TFBS_cnc mE1_TFBS_HSA mE1_TFBS_sbb mE1_TFBS_Trl
3R	15358856	3R_15358856_SNP [dm3] 3R_19533134_SNP [dm6]	A	C	1.625e^-5	0.858	INTRON Dys
2R	20110836	2R_20110836_SNP [dm3] 2R_24223313_SNP [dm6]	A	C	1.642e^-5	0.858	INTRON CG30419
2R	20110872	2R_20110872_SNP [dm3] 2R_24223349_SNP [dm6]	T	C	1.687e^-5	0.858	INTRON CG30419
3L	14390146	3L_14390146_SNP [dm3] 3L_14397046_SNP [dm6]	A	T	1.691e^-5	0.858	INTERGENIC
2R	4938827	2R_4938827_SNP [dm3] 2R_9051322_SNP [dm6]	A	G	1.700e^-5	0.858	INTRON CG13743	TF binding site BDTNP1_TFBS_twi mE1_TFBS_cad mE1_TFBS_HSA
2R	19205379	2R_19205379_SNP [dm3] 2R_23317856_SNP [dm6]	A	C	1.738e^-5	0.858	DOWNSTREAM CG30412 INTERGENIC	TF binding site mE1_TFBS_chinmo
3L	3096086	3L_3096086_SNP [dm3] 3L_3096086_SNP [dm6]	C	T	1.785e^-5	0.858	UPSTREAM tRNA:CR32287 & 1 other DOWNSTREAM tRNA:CR32288 & 2 others INTRON Cht7	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_Med
2R	1342879	2R_1342879_SNP [dm3] 2R_5455374_SNP [dm6]	A	G	1.972e^-5	0.925	UTR_3_PRIME laccase2	TF binding site mE1_TFBS_HSA mE1_TFBS_sbb
3R	4602711	3R_4602711_SNP [dm3] 3R_8776989_SNP [dm6]	T	C	2.086e^-5	0.933	SYNONYMOUS_CODING Pif1B UTR_3_PRIME Pif1A	TF binding site mE1_TFBS_HSA
3L	18295686	3L_18295686_SNP [dm3] 3L_18302586_SNP [dm6]	C	T	2.122e^-5	0.933	DOWNSTREAM grim INTERGENIC	TF binding site BDTNP1_TFBS_twi
2L	1163165	2L_1163165_SNP [dm3] 2L_1163165_SNP [dm6]	T	A	2.176e^-5	0.933	UTR_5_PRIME CG5080 INTRON CG5080	TF binding site mE1_TFBS_cad
3R	9541367	3R_9541367_SNP [dm3] 3R_13715645_SNP [dm6]	T	A	2.189e^-5	0.933	UTR_5_PRIME CG34383	TF binding site BDTNP1_TFBS_bcd BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_gt BDTNP1_TFBS_hb BDTNP1_TFBS_Med BDTNP1_TFBS_prd BDTNP1_TFBS_shn BDTNP1_TFBS_slp1 BDTNP1_TFBS_sna BDTNP1_TFBS_twi BDTNP1_TFBS_z mE1_TFBS_cad mE1_TFBS_D mE1_TFBS_disco mE1_TFBS_hkb mE1_TFBS_HSA mE1_TFBS_sens mE1_TFBS_Trl
3L	21044853	3L_21044853_SNP [dm3] 3L_21051753_SNP [dm6]	C	T	2.423e^-5	1	INTRON siz
3L	3096495	3L_3096495_SNP [dm3] 3L_3096495_SNP [dm6]	T	C	2.691e^-5	1	UPSTREAM tRNA:CR32285 DOWNSTREAM tRNA:CR32288 & 3 others INTRON Cht7	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_Med mE1_TFBS_HSA
3L	14390168	3L_14390168_SNP [dm3] 3L_14397068_SNP [dm6]	T	A	2.709e^-5	1	INTERGENIC
3L	13939838	3L_13939838_INS [dm3] 3L_13946738_INS [dm6]	T	TT	2.722e^-5	1	INTRON CG32137
3R	4168820	3R_4168820_SNP [dm3] 3R_8343098_SNP [dm6]	G	C	2.749e^-5	1	SYNONYMOUS_CODING CG9601 UPSTREAM CG18193	TF binding site BDTNP1_TFBS_dl mE1_TFBS_HSA mE1_TFBS_sens
2R	3485607	2R_3485607_SNP [dm3] 2R_7598102_SNP [dm6]	T	G	2.939e^-5	1	INTRON CG1358	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_twi mE1_TFBS_HSA
3L	12325453	3L_12325453_SNP [dm3] 3L_12332353_SNP [dm6]	A	G	3.033e^-5	1	INTRON CG32105
2R	1342866	2R_1342866_SNP [dm3] 2R_5455361_SNP [dm6]	A	G	3.148e^-5	1	UTR_3_PRIME laccase2	TF binding site mE1_TFBS_HSA mE1_TFBS_sbb
X	13692213	X_13692213_SNP [dm3] X_13798180_SNP [dm6]	T	A	3.177e^-5	1	UPSTREAM CG10996 INTRON inaE
2R	15793091	2R_15793091_SNP [dm3] 2R_19905586_SNP [dm6]	C	G	3.333e^-5	1	INTERGENIC
3L	12199282	3L_12199282_SNP [dm3] 3L_12206182_SNP [dm6]	T	A	3.345e^-5	1	UPSTREAM CG5626 DOWNSTREAM app INTERGENIC	TF binding site mE1_TFBS_HSA mE1_TFBS_zfh1
3R	13150606	3R_13150606_SNP [dm3] 3R_17324884_SNP [dm6]	G	A	3.355e^-5	1	INTRON beat-IIa
2L	13123672	2L_13123672_SNP [dm3] 2L_13123672_SNP [dm6]	A	T	3.401e^-5	1	INTERGENIC
3R	16395426	3R_16395426_SNP [dm3] 3R_20569704_SNP [dm6]	G	A	3.450e^-5	1	INTRON CG5191
3R	15358802	3R_15358802_SNP [dm3] 3R_19533080_SNP [dm6]	C	A	3.479e^-5	1	INTRON Dys
3L	18493976	3L_18493976_SNP [dm3] 3L_18500876_SNP [dm6]	C	A	3.550e^-5	1	INTRON AstC-R2
2L	10204158	2L_10204158_SNP [dm3] 2L_10204158_SNP [dm6]	T	C	3.575e^-5	1	DOWNSTREAM CG5734 INTRON CG34159	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_Med mE1_TFBS_HSA
3L	2010863	3L_2010863_SNP [dm3] 3L_2010863_SNP [dm6]	T	C	3.606e^-5	1	UPSTREAM CR44535 INTERGENIC
3L	11601321	3L_11601321_SNP [dm3] 3L_11608221_SNP [dm6]	T	C	3.614e^-5	1	UTR_5_PRIME CG6091 INTRON CG6091	TF binding site mE1_TFBS_HSA mE1_TFBS_sens
3L	11601563	3L_11601563_SNP [dm3] 3L_11608463_SNP [dm6]	G	T	3.751e^-5	1	UPSTREAM CG6091 INTRON CG6091	TF binding site BDTNP1_TFBS_dl mE1_TFBS_HSA
2R	16758224	2R_16758224_SNP [dm3] 2R_20870719_SNP [dm6]	T	A	3.759e^-5	1	UPSTREAM Ipk1 DOWNSTREAM IM14 INTERGENIC	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_twi mE1_TFBS_bab1 mE1_TFBS_disco mE1_TFBS_GATAe mE1_TFBS_HSA mE1_TFBS_jumu mE1_TFBS_Ubx
2L	2400167	2L_2400167_SNP [dm3] 2L_2400167_SNP [dm6]	C	T	3.858e^-5	1	UPSTREAM VGlut INTRON VGlut
3R	10353207	3R_10353207_SNP [dm3] 3R_14527485_SNP [dm6]	G	T	4.025e^-5	1	INTRON Pde6
3R	15358848	3R_15358848_SNP [dm3] 3R_19533126_SNP [dm6]	G	A	4.154e^-5	1	INTRON Dys
3R	13201768	3R_13201768_SNP [dm3] 3R_17376046_SNP [dm6]	T	C	4.364e^-5	1	SYNONYMOUS_CODING Brf
2L	10209192	2L_10209192_SNP [dm3] 2L_10209192_SNP [dm6]	G	C	4.397e^-5	1	DOWNSTREAM CG5727 INTRON CG5731 EXON CR44845	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_hb BDTNP1_TFBS_Med mE1_TFBS_D mE1_TFBS_HSA mE1_TFBS_Trl
2L	10209194	2L_10209194_SNP [dm3] 2L_10209194_SNP [dm6]	T	C	4.412e^-5	1	DOWNSTREAM CG5727 INTRON CG5731 EXON CR44845	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_hb BDTNP1_TFBS_Med mE1_TFBS_D mE1_TFBS_HSA mE1_TFBS_Trl
2L	2399755	2L_2399755_SNP [dm3] 2L_2399755_SNP [dm6]	G	A	4.535e^-5	1	UPSTREAM VGlut INTRON VGlut
2R	6423030	2R_6423030_SNP [dm3] 2R_10535525_SNP [dm6]	A	G	4.572e^-5	1	UPSTREAM lola DOWNSTREAM snoRNA:lola-a INTRON lola	TF binding site BDTNP1_TFBS_bcd BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_gt BDTNP1_TFBS_hb BDTNP1_TFBS_Med BDTNP1_TFBS_prd BDTNP1_TFBS_sna BDTNP1_TFBS_twi BDTNP1_TFBS_z mE1_TFBS_D mE1_TFBS_eve mE1_TFBS_HSA mE1_TFBS_Trl
2R	15793112	2R_15793112_SNP [dm3] 2R_19905607_SNP [dm6]	A	T	4.586e^-5	1	INTERGENIC
3R	15358846	3R_15358846_SNP [dm3] 3R_19533124_SNP [dm6]	A	C	4.645e^-5	1	INTRON Dys
3L	14390212	3L_14390212_SNP [dm3] 3L_14397112_SNP [dm6]	T	C	4.732e^-5	1	INTERGENIC	TF binding site BDTNP1_TFBS_dl
3L	12336230	3L_12336230_SNP [dm3] 3L_12343130_SNP [dm6]	T	C	4.758e^-5	1	INTRON CrzR
3R	5661448	3R_5661448_INS [dm3] 3R_9835726_INS [dm6]	C	CAA	4.787e^-5	1	INTRON Teh1
3R	15358858	3R_15358858_SNP [dm3] 3R_19533136_SNP [dm6]	G	T	4.822e^-5	1	INTRON Dys
2L	10230656	2L_10230656_SNP [dm3] 2L_10230656_SNP [dm6]	G	C	4.998e^-5	1	INTRON CG5694	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_Med BDTNP1_TFBS_twi mE1_TFBS_cad mE1_TFBS_D mE1_TFBS_HSA
3L	12199245	3L_12199245_SNP [dm3] 3L_12206145_SNP [dm6]	G	A	5.010e^-5	1	UPSTREAM CG5626 DOWNSTREAM app INTERGENIC	TF binding site mE1_TFBS_HSA mE1_TFBS_zfh1
X	22362736	X_22362736_SNP [dm3] X_22960900_SNP [dm6]	C	A	5.024e^-5	1	UTR_3_PRIME CG40485 UPSTREAM CG40486 DOWNSTREAM CG40485
2R	7503959	2R_7503959_SNP [dm3] 2R_11616454_SNP [dm6]	A	G	5.207e^-5	1	UPSTREAM tou DOWNSTREAM Egm EXON CR43907	TF binding site BDTNP1_TFBS_bcd BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_gt BDTNP1_TFBS_hb BDTNP1_TFBS_Med BDTNP1_TFBS_prd BDTNP1_TFBS_slp1 BDTNP1_TFBS_sna BDTNP1_TFBS_twi BDTNP1_TFBS_z mE1_TFBS_cad mE1_TFBS_disco mE1_TFBS_h mE1_TFBS_HSA mE1_TFBS_inv mE1_TFBS_Trl
3R	13145460	3R_13145460_SNP [dm3] 3R_17319738_SNP [dm6]	C	T	5.220e^-5	1	DOWNSTREAM beat-IIa INTERGENIC
X	18410446	X_18410446_SNP [dm3] X_18516413_SNP [dm6]	A	G	5.253e^-5	1	INTRON Aats-his & 1 other	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_prd BDTNP1_TFBS_twi mE1_TFBS_hkb mE1_TFBS_HSA mE1_TFBS_jumu mE1_TFBS_run mE1_TFBS_Trl
2L	20343221	2L_20343221_SNP [dm3] 2L_20343221_SNP [dm6]	A	T	5.561e^-5	1	INTRON spir
3R	12557224	3R_12557224_SNP [dm3] 3R_16731502_SNP [dm6]	T	G	5.581e^-5	1	INTRON Ubx	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_ftz BDTNP1_TFBS_gt BDTNP1_TFBS_Med BDTNP1_TFBS_prd BDTNP1_TFBS_sna BDTNP1_TFBS_twi mE1_TFBS_cad mE1_TFBS_HSA
X	9498848	X_9498848_SNP [dm3] X_9604815_SNP [dm6]	G	A	5.594e^-5	1	UTR_3_PRIME Lst8 DOWNSTREAM Gga
X	9498923	X_9498923_SNP [dm3] X_9604890_SNP [dm6]	G	A	5.594e^-5	1	SYNONYMOUS_CODING Lst8 DOWNSTREAM Gga
3R	25085289	3R_25085289_SNP [dm3] 3R_29259567_SNP [dm6]	G	C	5.688e^-5	1	INTERGENIC	regulatory region REDfly_CRMs TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_twi
3R	25085292	3R_25085292_SNP [dm3] 3R_29259570_SNP [dm6]	G	C	5.688e^-5	1	INTERGENIC	regulatory region REDfly_CRMs TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_twi
3R	15358800	3R_15358800_SNP [dm3] 3R_19533078_SNP [dm6]	A	C	5.831e^-5	1	INTRON Dys
3L	1132346	3L_1132346_SNP [dm3] 3L_1132346_SNP [dm6]	T	A	5.923e^-5	1	INTERGENIC	regulatory region REDfly_CRMs
3L	13396255	3L_13396255_SNP [dm3] 3L_13403155_SNP [dm6]	T	G	5.924e^-5	1	INTERGENIC	regulatory region REDfly_CRMs silencer mE1_HDAC_PRE TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_hb BDTNP1_TFBS_Med BDTNP1_TFBS_sna BDTNP1_TFBS_twi mE1_TFBS_bab1 mE1_TFBS_cad mE1_TFBS_HSA mE1_TFBS_sens
3R	5948642	3R_5948642_SNP [dm3] 3R_10122920_SNP [dm6]	C	A	5.931e^-5	1	SYNONYMOUS_CODING knk DOWNSTREAM CG31477	TF binding site mE1_TFBS_HSA
3R	13150611	3R_13150611_SNP [dm3] 3R_17324889_SNP [dm6]	C	G	5.945e^-5	1	INTRON beat-IIa
2R	9126244	2R_9126244_SNP [dm3] 2R_13238739_SNP [dm6]	A	G	5.975e^-5	1	SYNONYMOUS_CODING CG4679 UPSTREAM GstE14	TF binding site mE1_TFBS_cad mE1_TFBS_sens
X	22362548	X_22362548_SNP [dm3] X_22960712_SNP [dm6]	A	T	6.009e^-5	1	UTR_3_PRIME CG40485 UPSTREAM CG40486 DOWNSTREAM CG40485
3R	12557274	3R_12557274_SNP [dm3] 3R_16731552_SNP [dm6]	G	A	6.047e^-5	1	INTRON Ubx	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_ftz BDTNP1_TFBS_gt BDTNP1_TFBS_Med BDTNP1_TFBS_prd BDTNP1_TFBS_sna BDTNP1_TFBS_twi mE1_TFBS_cad mE1_TFBS_HSA
3R	21161330	3R_21161330_SNP [dm3] 3R_25335608_SNP [dm6]	A	G	6.112e^-5	1	SYNONYMOUS_CODING CG11902 UPSTREAM CG10669	TF binding site mE1_TFBS_HSA mE1_TFBS_sens mE1_TFBS_Ubx

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