DGRPool

GWAS results were computed on dm3 genome.

If you want to make an independent analysis with DGRP2 or PLINK2, you can find here prepared input files for this phenotype: DGRP2 results PLINK2 results

Full results

Phenotype distribution and association to known covariates

Shapiro-Wilk test of normality (p-value=1.426e^-1)
Null hypothesis is not rejected (p>0.05).
Interpretation: NORMAL distribution of this phenotype

We tested for association of this phenotype with 6 known covariates Huang et al., 2014 using both ANOVA and Kruskal-Wallis association tests.

ANOVA test

Covariate	p-value	Significance
In_2L_t	0.07973

Kruskal-Wallis test
No significant result.

PNG PDF

QQplot

PNG PDF

Manhattan plot

GWAS results

890 significant hits at p-value < 0.001.

We performed a gene enrichment calculation (Fisher's exact test) of all GWAS-associated genes at p < 0.001. Genesets were taken from two database: the Gene Ontology (GO) and the FlyBase phenotypes. GO enrichment FlyBase phenotype enrichment

Variant impact HIGH MODERATE LOW MODIFIER

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Chromosome	Position	ID	Reference	Variant	p-value	FDR_BH	Transcript annotation	Binding site annotation
2R	17722253	2R_17722253_SNP [dm3] 2R_21834748_SNP [dm6]	G	A	4.200e^-6	0.86	SYNONYMOUS_CODING CG4363 DOWNSTREAM CG4377
X	11111178	X_11111178_SNP [dm3] X_11217145_SNP [dm6]	T	C	1.225e^-5	0.86	INTERGENIC
2R	17436228	2R_17436228_SNP [dm3] 2R_21548723_SNP [dm6]	C	A	1.548e^-5	0.86	INTRON Egfr	TF binding site BDTNP1_TFBS_dl
2R	8911925	2R_8911925_SNP [dm3] 2R_13024420_SNP [dm6]	A	G	1.571e^-5	0.86	INTERGENIC
3L	18492202	3L_18492202_SNP [dm3] 3L_18499102_SNP [dm6]	G	A	1.571e^-5	0.86	INTRON AstC-R2
3L	9620585	3L_9620585_SNP [dm3] 3L_9627485_SNP [dm6]	A	T	1.860e^-5	0.86	DOWNSTREAM CG3335 INTRON Or67b
3L	9620726	3L_9620726_SNP [dm3] 3L_9627626_SNP [dm6]	C	T	2.050e^-5	0.86	NON_SYNONYMOUS_CODING Or67b DOWNSTREAM CG3335
3L	9620811	3L_9620811_SNP [dm3] 3L_9627711_SNP [dm6]	G	A	2.050e^-5	0.86	UPSTREAM CG8336 DOWNSTREAM CG3335 INTRON Or67b
3L	9620816	3L_9620816_SNP [dm3] 3L_9627716_SNP [dm6]	T	A	2.050e^-5	0.86	UPSTREAM CG8336 DOWNSTREAM CG3335 INTRON Or67b
3L	9621119	3L_9621119_SNP [dm3] 3L_9628019_SNP [dm6]	T	A	2.050e^-5	0.86	UPSTREAM CG8336 DOWNSTREAM CG3335 INTRON Or67b
3R	25356335	3R_25356335_SNP [dm3] 3R_29530613_SNP [dm6]	T	A	2.363e^-5	0.86	INTERGENIC
3L	9620493	3L_9620493_SNP [dm3] 3L_9627393_SNP [dm6]	C	A	2.524e^-5	0.86	UTR_3_PRIME Or67b DOWNSTREAM CG3335
3L	9621095	3L_9621095_SNP [dm3] 3L_9627995_SNP [dm6]	A	G	2.524e^-5	0.86	SYNONYMOUS_CODING Or67b UPSTREAM CG8336 DOWNSTREAM CG3335
3R	918518	3R_918518_SNP [dm3] 3R_5092796_SNP [dm6]	G	T	2.524e^-5	0.86	INTERGENIC
3R	24872160	3R_24872160_SNP [dm3] 3R_29046438_SNP [dm6]	A	C	2.996e^-5	0.86	UTR_3_PRIME Pkc98E
2R	5427749	2R_5427749_SNP [dm3] 2R_9540244_SNP [dm6]	G	A	3.053e^-5	0.86	UTR_3_PRIME brp	TF binding site BDTNP1_TFBS_dl mE1_TFBS_HSA
2R	11653566	2R_11653566_SNP [dm3] 2R_15766061_SNP [dm6]	T	G	3.183e^-5	0.86	SYNONYMOUS_CODING Zasp52 INTRON Zasp52	TF binding site mE1_TFBS_cad mE1_TFBS_HSA mE1_TFBS_sens
X	18375851	X_18375851_SNP [dm3] X_18481818_SNP [dm6]	C	T	4.192e^-5	0.86	SYNONYMOUS_CODING Hayan DOWNSTREAM CR45714 INTRON Hayan	TF binding site mE1_TFBS_HSA mE1_TFBS_inv
X	11111129	X_11111129_SNP [dm3] X_11217096_SNP [dm6]	T	C	4.318e^-5	0.86	INTERGENIC
2R	11415155	2R_11415155_SNP [dm3] 2R_15527650_SNP [dm6]	C	T	4.507e^-5	0.86	SYNONYMOUS_CODING Khc-73	TF binding site BDTNP1_TFBS_dl
2R	11416422	2R_11416422_SNP [dm3] 2R_15528917_SNP [dm6]	T	C	4.507e^-5	0.86	INTRON Khc-73	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_prd BDTNP1_TFBS_twi mE1_TFBS_HSA
3L	4978489	3L_4978489_SNP [dm3] 3L_4978489_SNP [dm6]	A	G	4.730e^-5	0.86	INTRON Con
2L	4794246	2L_4794246_SNP [dm3] 2L_4794246_SNP [dm6]	A	C	4.780e^-5	0.86	UPSTREAM CG3294 & 1 other INTERGENIC	TF binding site BDTNP1_TFBS_dl
2L	21916487	2L_21916487_SNP [dm3] 2L_21916487_SNP [dm6]	T	A	4.839e^-5	0.86	NON_SYNONYMOUS_CODING CG11634
2R	17025724	2R_17025724_SNP [dm3] 2R_21138219_SNP [dm6]	G	C	5.211e^-5	0.86	SPLICE_SITE_REGION Magi INTRON Magi	TF binding site mE1_TFBS_chinmo
2R	8911929	2R_8911929_SNP [dm3] 2R_13024424_SNP [dm6]	C	A	5.308e^-5	0.86	INTERGENIC
3R	15327713	3R_15327713_SNP [dm3] 3R_19501991_SNP [dm6]	G	T	5.308e^-5	0.86	INTRON Dys
3R	18328932	3R_18328932_SNP [dm3] 3R_22503210_SNP [dm6]	T	A	5.308e^-5	0.86	INTRON Octbeta1R
3R	25265193	3R_25265193_SNP [dm3] 3R_29439471_SNP [dm6]	A	T	5.308e^-5	0.86	INTRON Ptp99A
3L	5820932	3L_5820932_SNP [dm3] 3L_5827832_SNP [dm6]	A	T	5.580e^-5	0.86	INTRON vn
3L	9620785	3L_9620785_SNP [dm3] 3L_9627685_SNP [dm6]	A	G	5.724e^-5	0.86	SPLICE_SITE_REGION Or67b UPSTREAM CG8336 DOWNSTREAM CG3335 INTRON Or67b
3L	4978443	3L_4978443_SNP [dm3] 3L_4978443_SNP [dm6]	T	G	5.910e^-5	0.86	INTRON Con
3L	14527380	3L_14527380_SNP [dm3] 3L_14534280_SNP [dm6]	G	C	5.952e^-5	0.86	INTRON bbg	TF binding site mE1_TFBS_Trl
2L	9011520	2L_9011520_SNP [dm3] 2L_9011520_SNP [dm6]	A	T	5.962e^-5	0.86	UPSTREAM CG13102 DOWNSTREAM CG9568 INTERGENIC
2L	9011522	2L_9011522_DEL [dm3] 2L_9011522_DEL [dm6]	AAA	A	5.962e^-5	0.86	UPSTREAM CG13102 DOWNSTREAM CG9568 INTERGENIC
2L	6566375	2L_6566375_SNP [dm3] 2L_6566375_SNP [dm6]	G	C	6.458e^-5	0.86	UPSTREAM CG13766 INTERGENIC
X	5575950	X_5575950_SNP [dm3] X_5681917_SNP [dm6]	A	C	6.924e^-5	0.86	SYNONYMOUS_CODING Gr5a EXON CR44958	TF binding site mE1_TFBS_cad mE1_TFBS_HSA
2L	7189211	2L_7189211_SNP [dm3] 2L_7189211_SNP [dm6]	G	C	7.124e^-5	0.86	INTRON CG4496	TF binding site BDTNP1_TFBS_dl mE1_TFBS_cad mE1_TFBS_HSA
3R	18704146	3R_18704146_SNP [dm3] 3R_22878424_SNP [dm6]	T	G	8.221e^-5	0.86	INTERGENIC	TF binding site BDTNP1_TFBS_gt BDTNP1_TFBS_Med BDTNP1_TFBS_twi mE1_TFBS_cad
2L	17430732	2L_17430732_SNP [dm3] 2L_17430732_SNP [dm6]	G	A	8.342e^-5	0.86	DOWNSTREAM CR44409 INTRON Dif
X	11691832	X_11691832_INS [dm3] X_11797799_INS [dm6]	G	GA	8.577e^-5	0.86	UTR_3_PRIME CkIIbeta DOWNSTREAM CkIIbeta INTRON CkIIbeta	TF binding site BDTNP1_TFBS_dl mE1_TFBS_HSA
3L	7956155	3L_7956155_SNP [dm3] 3L_7963055_SNP [dm6]	C	A	8.638e^-5	0.86	INTERGENIC	TF binding site mE1_TFBS_cad mE1_TFBS_HSA
2L	6336150	2L_6336150_SNP [dm3] 2L_6336150_SNP [dm6]	A	G	8.647e^-5	0.86	SYNONYMOUS_CODING Pez	TF binding site mE1_TFBS_cad mE1_TFBS_chinmo mE1_TFBS_HSA
2L	12323240	2L_12323240_SNP [dm3] 2L_12323240_SNP [dm6]	A	G	9.065e^-5	0.86	INTERGENIC
3L	1885006	3L_1885006_SNP [dm3] 3L_1885006_SNP [dm6]	G	T	9.456e^-5	0.86	UTR_5_PRIME Aprt UPSTREAM Tmhs & 1 other	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_Med BDTNP1_TFBS_twi mE1_TFBS_HSA mE1_TFBS_Trl
3L	1885017	3L_1885017_SNP [dm3] 3L_1885017_SNP [dm6]	G	T	9.456e^-5	0.86	UTR_5_PRIME Aprt UPSTREAM Tmhs & 1 other	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_Med BDTNP1_TFBS_twi mE1_TFBS_HSA mE1_TFBS_Trl
3L	1885018	3L_1885018_SNP [dm3] 3L_1885018_SNP [dm6]	G	T	9.456e^-5	0.86	UTR_5_PRIME Aprt UPSTREAM Tmhs & 1 other	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_Med BDTNP1_TFBS_twi mE1_TFBS_HSA mE1_TFBS_Trl
3L	20623215	3L_20623215_SNP [dm3] 3L_20630115_SNP [dm6]	T	C	9.456e^-5	0.86	INTERGENIC	TF binding site BDTNP1_TFBS_bcd BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_hb BDTNP1_TFBS_Med BDTNP1_TFBS_prd BDTNP1_TFBS_slp1 BDTNP1_TFBS_sna BDTNP1_TFBS_twi mE1_TFBS_HSA mE1_TFBS_sens
2R	14939481	2R_14939481_SNP [dm3] 2R_19051976_SNP [dm6]	C	T	9.681e^-5	0.86	INTERGENIC
X	6388208	X_6388208_SNP [dm3] X_6494175_SNP [dm6]	G	A	9.681e^-5	0.86	INTRON CG42340
3R	20823341	3R_20823341_SNP [dm3] 3R_24997619_SNP [dm6]	G	A	9.856e^-5	0.86	INTRON tnc	TF binding site mE1_TFBS_cad mE1_TFBS_chinmo
X	20391131	X_20391131_SNP [dm3] X_20520104_SNP [dm6]	T	G	9.856e^-5	0.86	INTRON Tak1
X	8940282	X_8940282_SNP [dm3] X_9046249_SNP [dm6]	T	G	9.900e^-5	0.86	INTRON CR43836 & 1 other
3R	22890521	3R_22890521_DEL [dm3] 3R_27064799_DEL [dm6]	AA	C	9.911e^-5	0.86	UPSTREAM Nep5 & 1 other DOWNSTREAM spz INTERGENIC
X	9202607	X_9202607_SNP [dm3] X_9308574_SNP [dm6]	G	T	9.968e^-5	0.86	INTERGENIC
3L	12432091	3L_12432091_SNP [dm3] 3L_12438991_SNP [dm6]	G	A	1.021e^-4	0.86	INTERGENIC	TF binding site BDTNP1_TFBS_dl mE1_TFBS_HSA
3L	16185639	3L_16185639_SNP [dm3] 3L_16192539_SNP [dm6]	C	A	1.021e^-4	0.86	INTERGENIC
2R	5427731	2R_5427731_SNP [dm3] 2R_9540226_SNP [dm6]	G	T	1.041e^-4	0.86	UTR_3_PRIME brp	TF binding site BDTNP1_TFBS_dl mE1_TFBS_HSA
3L	1884972	3L_1884972_SNP [dm3] 3L_1884972_SNP [dm6]	C	T	1.049e^-4	0.86	UTR_5_PRIME Aprt UPSTREAM Tmhs & 1 other	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_Med BDTNP1_TFBS_twi mE1_TFBS_HSA mE1_TFBS_Trl
3L	16190193	3L_16190193_SNP [dm3] 3L_16197093_SNP [dm6]	T	A	1.052e^-4	0.86	UPSTREAM CG33687 EXON CR45437
3R	921448	3R_921448_SNP [dm3] 3R_5095726_SNP [dm6]	A	G	1.071e^-4	0.86	INTERGENIC	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_prd mE1_TFBS_HSA
3R	9839233	3R_9839233_SNP [dm3] 3R_14013511_SNP [dm6]	A	C	1.084e^-4	0.86	INTRON rdx	TF binding site BDTNP1_TFBS_dl BDTNP1_TFBS_twi mE1_TFBS_HSA
2R	4135877	2R_4135877_SNP [dm3] 2R_8248372_SNP [dm6]	A	C	1.134e^-4	0.86	DOWNSTREAM CG44439 INTERGENIC	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_ftz BDTNP1_TFBS_gt BDTNP1_TFBS_hb BDTNP1_TFBS_kni BDTNP1_TFBS_Med BDTNP1_TFBS_prd BDTNP1_TFBS_shn BDTNP1_TFBS_slp1 BDTNP1_TFBS_sna BDTNP1_TFBS_tll BDTNP1_TFBS_twi mE1_TFBS_cad mE1_TFBS_D mE1_TFBS_disco mE1_TFBS_eve mE1_TFBS_hkb mE1_TFBS_HSA mE1_TFBS_sens
3L	4978279	3L_4978279_SNP [dm3] 3L_4978279_SNP [dm6]	C	T	1.146e^-4	0.86	INTRON Con
2L	7168682	2L_7168682_SNP [dm3] 2L_7168682_SNP [dm6]	A	C	1.147e^-4	0.86	INTERGENIC
2R	7403149	2R_7403149_SNP [dm3] 2R_11515644_SNP [dm6]	C	T	1.170e^-4	0.86	INTERGENIC	TF binding site BDTNP1_TFBS_twi
3L	13941266	3L_13941266_SNP [dm3] 3L_13948166_SNP [dm6]	G	T	1.170e^-4	0.86	INTRON CG32137
3L	8124936	3L_8124936_SNP [dm3] 3L_8131836_SNP [dm6]	T	C	1.170e^-4	0.86	SYNONYMOUS_CODING CG7504 DOWNSTREAM CdsA
3L	8124987	3L_8124987_SNP [dm3] 3L_8131887_SNP [dm6]	T	A	1.170e^-4	0.86	NON_SYNONYMOUS_CODING CG7504 DOWNSTREAM CdsA
2L	6559501	2L_6559501_SNP [dm3] 2L_6559501_SNP [dm6]	C	T	1.185e^-4	0.86	UTR_5_PRIME CG11043 UTR_3_PRIME CG9596 DOWNSTREAM CG9596 & 1 other	TF binding site mE1_TFBS_cad
2L	3750983	2L_3750983_SNP [dm3] 2L_3750983_SNP [dm6]	C	G	1.196e^-4	0.86	INTRON CG10019
3R	22848917	3R_22848917_SNP [dm3] 3R_27023195_SNP [dm6]	C	T	1.206e^-4	0.86	UTR_5_PRIME CG6277 INTRON RYa-R
3R	22848918	3R_22848918_SNP [dm3] 3R_27023196_SNP [dm6]	G	A	1.206e^-4	0.86	START_GAINED CG6277 UTR_5_PRIME CG6277 INTRON RYa-R
2L	6259241	2L_6259241_SNP [dm3] 2L_6259241_SNP [dm6]	G	A	1.212e^-4	0.86	INTRON Ddr	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_Med BDTNP1_TFBS_prd BDTNP1_TFBS_twi mE1_TFBS_HSA
3L	12050524	3L_12050524_SNP [dm3] 3L_12057424_SNP [dm6]	T	A	1.212e^-4	0.86	UPSTREAM rols INTRON rols	TF binding site mE1_TFBS_HSA mE1_TFBS_sens
2L	580737	2L_580737_SNP [dm3] 2L_580737_SNP [dm6]	C	T	1.237e^-4	0.86	INTERGENIC
2L	6340794	2L_6340794_INS [dm3] 2L_6340794_INS [dm6]	A	AA	1.237e^-4	0.86	UPSTREAM Cpr INTRON Cpr	TF binding site mE1_TFBS_cnc mE1_TFBS_zfh1
2L	6340799	2L_6340799_INS [dm3] 2L_6340799_INS [dm6]	T	TTGTTAGTTTT...	1.237e^-4	0.86	UPSTREAM Cpr INTRON Cpr	TF binding site mE1_TFBS_cnc mE1_TFBS_zfh1
2L	6340809	2L_6340809_SNP [dm3] 2L_6340809_SNP [dm6]	C	T	1.237e^-4	0.86	UPSTREAM Cpr INTRON Cpr	TF binding site mE1_TFBS_cnc mE1_TFBS_zfh1
3R	19804622	3R_19804622_SNP [dm3] 3R_23978900_SNP [dm6]	G	A	1.304e^-4	0.86	INTRON CG31140
3L	20616024	3L_20616024_SNP [dm3] 3L_20622924_SNP [dm6]	G	A	1.314e^-4	0.86	DOWNSTREAM CR44684 INTERGENIC	TF binding site BDTNP1_TFBS_bcd BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_gt BDTNP1_TFBS_hb BDTNP1_TFBS_prd BDTNP1_TFBS_tll BDTNP1_TFBS_z mE1_TFBS_cad mE1_TFBS_HSA mE1_TFBS_Kr mE1_TFBS_Trl
2L	19614904	2L_19614904_SNP [dm3] 2L_19614904_SNP [dm6]	C	T	1.330e^-4	0.86	INTRON Lar
3L	3297604	3L_3297604_SNP [dm3] 3L_3297604_SNP [dm6]	C	T	1.335e^-4	0.86	SYNONYMOUS_CODING CG12009
2R	11844909	2R_11844909_SNP [dm3] 2R_15957404_SNP [dm6]	A	G	1.346e^-4	0.86	UPSTREAM Strn-Mlck DOWNSTREAM CG8366 INTRON Strn-Mlck	TF binding site mE1_TFBS_cad
2R	11844926	2R_11844926_SNP [dm3] 2R_15957421_SNP [dm6]	T	C	1.346e^-4	0.86	UPSTREAM Strn-Mlck DOWNSTREAM CG8366 INTRON Strn-Mlck	TF binding site mE1_TFBS_cad
2R	5444062	2R_5444062_SNP [dm3] 2R_9556557_SNP [dm6]	T	G	1.346e^-4	0.86	INTERGENIC
2R	15314362	2R_15314362_SNP [dm3] 2R_19426857_SNP [dm6]	T	A	1.351e^-4	0.86	INTRON CalpA
2R	8433622	2R_8433622_SNP [dm3] 2R_12546117_SNP [dm6]	C	A	1.351e^-4	0.86	INTRON fra & 1 other
3L	3671054	3L_3671054_SNP [dm3] 3L_3671054_SNP [dm6]	T	A	1.351e^-4	0.86	INTERGENIC
3L	7307032	3L_7307032_SNP [dm3] 3L_7313932_SNP [dm6]	G	A	1.351e^-4	0.86	INTERGENIC
3L	1409783	3L_1409783_SNP [dm3] 3L_1409783_SNP [dm6]	A	G	1.364e^-4	0.86	INTRON Ptp61F	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_ftz BDTNP1_TFBS_Med BDTNP1_TFBS_prd BDTNP1_TFBS_twi mE1_TFBS_cad mE1_TFBS_chinmo mE1_TFBS_HSA
2R	13907150	2R_13907150_SNP [dm3] 2R_18019645_SNP [dm6]	A	C	1.370e^-4	0.86	SYNONYMOUS_CODING CG5084 UPSTREAM CG10910	TF binding site mE1_TFBS_cad
2R	10986649	2R_10986649_DEL [dm3] 2R_15099144_DEL [dm6]	CTATG	C	1.381e^-4	0.86	INTERGENIC	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_Med BDTNP1_TFBS_sna BDTNP1_TFBS_twi
2R	10986668	2R_10986668_SNP [dm3] 2R_15099163_SNP [dm6]	T	G	1.381e^-4	0.86	INTERGENIC	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_Med BDTNP1_TFBS_sna BDTNP1_TFBS_twi
2R	10986681	2R_10986681_INS [dm3] 2R_15099176_INS [dm6]	A	TCCTTAATTTCC...	1.381e^-4	0.86	INTERGENIC	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_Med BDTNP1_TFBS_sna BDTNP1_TFBS_twi
2R	10986689	2R_10986689_SNP [dm3] 2R_15099184_SNP [dm6]	A	T	1.381e^-4	0.86	INTERGENIC	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_Med BDTNP1_TFBS_sna BDTNP1_TFBS_twi
2R	10986690	2R_10986690_SNP [dm3] 2R_15099185_SNP [dm6]	A	T	1.381e^-4	0.86	INTERGENIC	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_Med BDTNP1_TFBS_sna BDTNP1_TFBS_twi
3L	16257560	3L_16257560_SNP [dm3] 3L_16264460_SNP [dm6]	G	T	1.401e^-4	0.86	UPSTREAM CG13069 DOWNSTREAM CG13051 & 1 other INTERGENIC	TF binding site mE1_TFBS_inv
3R	11400179	3R_11400179_SNP [dm3] 3R_15574457_SNP [dm6]	C	T	1.411e^-4	0.86	INTERGENIC	TF binding site mE1_TFBS_chinmo mE1_TFBS_inv
2R	10986730	2R_10986730_SNP [dm3] 2R_15099225_SNP [dm6]	A	T	1.445e^-4	0.86	INTERGENIC	TF binding site BDTNP1_TFBS_da BDTNP1_TFBS_dl BDTNP1_TFBS_Med BDTNP1_TFBS_sna BDTNP1_TFBS_twi

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By genomic position
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LogMiu60 phenotype

Phenotype distribution and association to known covariates

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GWAS results